Moscow in March accused Washington of funding research into the development of biological weapons in Ukraine, which has invaded by Russia nearly five months ago.

Washington and Kyiv denied the existence of laboratories intended to produce biological weapons in Ukraine, with the United States saying the allegations were a sign that Moscow may use such tactics itself.

Izumi Nakamitsu, the UN's Under-Secretary-General of Disarmament Affairs, also said in March that the UN "was not aware of any biological weapons program in Ukraine".

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