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Khamenei repeated the accusations in a tweet that was removed by Twitter along with a message saying it violated the platform’s rules against misinformation.

Iran launched human trials of its first domestic COVID-19 vaccine candidate late last month, saying it could help Iran defeat the pandemic despite U.S. sanctions that affect its ability to import vaccines.

Tensions between Washington and Tehran have risen since 2018, when president Donald Trump abandoned a 2015 nuclear deal and reimposed sanctions to pressure Iran into negotiating stricter curbs on its nuclear program, ballistic missile development and support for regional proxy forces.

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In retaliation for US sanctions, which were lifted under the nuclear deal, Tehran has gradually violated the accord. US president-elect Joe Biden, who takes office on 20 January, has pledged to rejoin the agreement if Tehran also returns to full compliance.

Khamenei said Tehran was in no rush for the United States to re-enter the deal, but that sanctions on the Islamic Republic must be lifted immediately.

Iran’s utmost authority, Khamenei ruled out any talks over Tehran’s missile programme and Iran’s involvement in the Middle East, as demanded by the United States and some other major powers.

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“Contrary to the US, Iran’s involvement in the region creates stability and is aimed at preventing instability ... Iran’s involvement in the region is definite and will continue.”

Shortly before Khamenei’s speech, Iran’s elite Revolutionary Guards unveiled an underground missile base at an undisclosed Gulf location.

The West sees Iran’s missiles both as a conventional military threat to regional stability and a possible delivery mechanism for nuclear weapons should Tehran develop them.

But Iran, which has one of the biggest missile programmes in the Middle East, regards the programme as an important deterrent and retaliatory force against the United States and other adversaries - primarily Gulf Arabs - in the region in the event of war.

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